IMARCS Foundation

Module 26

Feeding and Nutrition

Why feeding matters, what animals eat, types of food, food size and variety, how much and how often, overfeeding and underfeeding, feeding time as a health check, feeding anemones, broadcast and target feeding a reef with food of many sizes, why corals need food as well as light, automatic feeding, live food, enrichment and storage, and finding the causes when feeding goes wrong.

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Key terms for this module

Read these first. Every one of them comes up in the sections below.

Herbivore An animal that eats algae and plants.
Carnivore An animal that eats other animals.
Omnivore An animal that eats both.
Planktivore An animal that picks small drifting food from the water.
Filter feeder An animal that strains tiny particles out of the water.
Phytoplankton Tiny drifting algae.
Rotifer A tiny animal, about 100 to 340 microns, grown as live food.
Artemia Brine shrimp, hatched and fed as live food.
Enrichment Feeding live prey a nutritious food just before the prey is fed out.
Micron One thousandth of a millimeter.
Broadcast feeding Putting food into the water for the whole system.
Target feeding Placing food on or near one animal.
Oral disc The flat top of an anemone, with the mouth at its center.
Polyp One coral animal: a mouth ringed by tentacles. A colony is many polyps.
Dosing pump A pump that adds a small, measured amount of liquid on a timer.

1. Why feeding matters

Food is how animals grow, keep their color and breed. It's also the biggest thing added to a system every day.

Where food goes Food goes in Eaten Growth, energy, color, breeding ! Not eaten Rots into ammonia, nitrate, phosphate ! Biggest input Food is the largest thing added to a system every day

Everything an animal is built from comes from what it eats: growth, energy, color, resistance to disease and the ability to breed. Good water can't make up for a poor diet.

Food is also the largest input to a system. Whatever goes in either becomes part of an animal or ends up as waste: uneaten food and droppings break down into ammonia, nitrate and phosphate (Module 8). So feeding is always two jobs at once: feeding the animals and managing what the food does to the water.

If Then
Food eaten Growth, energy, color, breeding.
Food not eaten Ammonia, nitrate, phosphate.

Check your understanding

A system's water tests are perfect, but its fish are thin and pale. What does that tell you?

Good water can't make up for a poor diet. Look at what's fed, how much, and who's eating.

Key point Feed the animals, manage the water.

2. What animals eat

Animals are built for different foods. The right food for one is the wrong food for another.

What animals eat Feeder Eats Herbivore Algae and seaweed, grazed all day Carnivore Meaty foods Omnivore Both meaty and plant foods Planktivore Small drifting food, often Filter feeder Tiny particles from the water Photosynthetic animals Clams, corals and anemones also get food from light

Herbivores, such as tangs, graze algae and seaweed through the day. Carnivores eat meaty foods. Omnivores, such as clownfish, eat both. Planktivores pick small drifting food from the water, a little at a time and often. Filter feeders, such as clams, strain tiny particles out of the water.

Clams, corals and anemones are also photosynthetic: algae living in their tissue turn light into food for them. Light isn't their only food, though, and they still feed. Before an animal is bought, find out what it eats (Module 24); a fish fed the wrong kind of food can starve with food in front of it.

If Then
Tang Herbivore: seaweed through the day.
Clam Light, plus filtered plankton.

Check your understanding

A tang gets mysis shrimp twice a day and is getting thin. Why?

It's a herbivore fed a carnivore's food. It needs algae and seaweed, through the day.

Key point Know what it eats before buying.

3. Types of food

Each kind of food has strengths and weaknesses. Variety covers the gaps.

Types of food Food Know this Dry pellets and flakes Easy to portion; loses vitamins Frozen Close to natural; keep it frozen Seaweed sheets For grazers, through the day Live Movement triggers feeding ! Variety No single food has everything

Dry pellets and flakes are easy to measure and store, and good ones are complete diets, but they lose vitamins once the container is opened. Frozen foods, such as mysis shrimp, are close to natural prey. Seaweed sheets feed grazers through the day. Live food moves, and the movement triggers feeding, which is why it works for newly arrived fish that refuse other food and for larvae.

No single food has everything. Rotating several foods through the week covers what any one of them lacks, and keeps animals taking a range of foods.

If Then
Only one food, every day Rotate several.
New fish refusing food Live food can start it.

Check your understanding

A newly arrived fish refuses pellets and frozen food for days. What might start it eating?

Live food: the movement triggers feeding. Then other foods are brought in.

Key point Variety covers the gaps.

4. Size and variety

Food has to fit the mouth. Different animals in one system need very different sizes.

Food size Phytoplankton Clams, rotifers, corals Rotifers 100 to 340 microns: larvae, corals Newly hatched Artemia Bigger larvae, small fish Pellets and frozen food Fish, anemones ! The mouth sets the limit Food has to fit to be eaten Many sizes A range of particle sizes feeds every level of the system

An animal's mouth sets the size of food it can eat. A fish that takes a pellet and spits it out may be telling you the pellet is too big. Newly hatched fish larvae can only take very small prey, such as rotifers, which measure about 100 to 340 microns. A micron is one thousandth of a millimeter.

A reef system holds mouths of every size: fish, corals, clams, worms, and the tiny animals living in the rock and sand. Feeding a range of particle sizes, from phytoplankton up to pellets, feeds every level of that food web, not only the fish.

If Then
Takes a pellet, spits it out May be too big.
Only large food fed Small mouths go hungry.

Check your understanding

A small fish rushes to the pellets, takes one, and spits it out, again and again. What do you check?

The size. The pellet may be too big for its mouth; offer a smaller food.

Key point Food has to fit the mouth.

5. How much and how often

Feed small amounts that get eaten, as often as the animal needs.

How much and how often Animal How often Most adult fish Small meals, once or twice a day Young and very active fish Several small meals a day Grazers Seaweed through the day ! Each meal Only what gets eaten within a few minutes

A meal is the amount that gets eaten within a few minutes. Most adult fish do well on small meals once or twice a day. Young fish and very active fish need several small meals a day. Grazers do best with seaweed available through the day.

Fish always act hungry, so begging is no guide. Watch what's actually eaten, and adjust: if food reaches the bottom and stays there, it was too much.

If Then
Food sitting on the bottom Too much.
Young, active fish Several small meals.

Check your understanding

Fish beg at the glass an hour after eating, so a coworker feeds again. What's wrong with that reasoning?

Fish always act hungry. The guide is what's eaten within a few minutes and whether any is left.

Key point What gets eaten in a few minutes.

6. Too much and too little

Both do harm. Overfeeding harms the water, and underfeeding harms the animals.

Too much and too little TOO MUCH TOO LITTLE ! Food left over ! It rots Ammonia and nutrients ! Algae And poor water Thin fish Pinched belly Slow growth Dull color More fighting Over what food there is ! Aim between them Enough for every animal, with nothing left to rot

Overfeeding is the most common feeding mistake. Uneaten food rots and releases ammonia, and nitrate and phosphate build up and feed algae. Ammonia, nitrate and phosphate aren't simply waste: at the right concentration they're food for corals, clams and algae, and at zero the animals that use them go hungry. The harm comes from too much.

Underfeeding shows on the animals: thin bodies, pinched bellies, slow growth, dull color and more fighting over what food there is. The answer to dirty water isn't starving the animals. Feed what gets eaten, and remove leftovers.

If Then
Algae and leftovers Overfeeding.
Thin, pinched, fighting Underfeeding.

Check your understanding

Algae is spreading, so a coworker stops feeding for a week. What's the risk, and the better answer?

The animals go hungry and get thin. Feed what gets eaten, remove leftovers, and deal with the excess nutrients.

Key point Enough for all, nothing left to rot.

7. Feeding time is a health check

Feeding is the best moment of the day to see every animal and notice what's changed.

Feeding time is a health check At feeding It may mean Comes out fast and eats Healthy Ignores food Often the first sign of trouble Takes food, spits it out Too big, wrong food, or sick One fish gets nothing Outcompeted Count heads at every meal Every animal, every time

A healthy fish comes out and eats. A fish that ignores food is often the first sign of trouble, before anything else shows. Some fish, clownfish among them, rest on the bottom and can look sick; when food goes in, a healthy one springs out and eats. Count heads at every meal.

Watch who gets the food. A bold fish can take everything while a shy or new one gets none. Feeding in two places at once, or placing food near the shy fish, makes sure every animal eats (Module 24).

If Then
Fish ignores food Often the first sign of trouble.
Shy fish gets none Feed in two places.

Check your understanding

A clownfish lies on the bottom and looks sick. How does feeding help you judge it?

A healthy one springs out and eats when food goes in. One that ignores food is a real warning.

Key point Count heads at every meal.

8. Feeding anemones

An anemone is fed by light and by what it catches. Fed anemones grow faster.

How to feed an anemone Raw, meaty marine food Fish, shrimp, mysis, scallop Cut smaller than the mouth Small pieces digest easily Flow off, place on the tentacles They grip it and carry it in Watch it swallow Keep fish and shrimp off it ! Never push food into the mouth The mouth tears easily. Let the anemone take it ! Spat out hours later The piece was too big, or the last meal was too recent

Anemones are carnivores. Their tentacles grip whatever lands on them and carry it to the mouth, at the center of the oral disc. The food is raw, meaty and marine: pieces of fish, shrimp, mysis, krill, scallop or clam, in variety. Cooked or seasoned food isn't used.

Cut each piece smaller than the anemone's mouth. Small pieces are digested more easily and are less likely to come back up. Turn the flow down or off so the food isn't blown away, then place the piece on the tentacles with tongs or a feeding tube and let go. A healthy anemone's tentacles feel sticky and hold the food, then fold it in toward the mouth. Never push food into the mouth: the mouth tears easily. Start a small or newly settled anemone on small pieces and a small amount, and increase as it keeps its meals down. A new anemone isn't fed until it has attached and settled.

Target feeding is commonly done once or twice a week. More food means faster growth, up to what the anemone can digest. In a heavily fed tank an anemone also catches food meant for the fish, and clownfish carry food to their own anemone: in field trials, anemones fed only by their clownfish grew about as well as anemones fed by hand. Fish and shrimp also steal, so watch until the meal is swallowed, or feed the fish first.

An anemone that spits food out hours later was given a piece that was too big, or was fed again too soon. Take the spat-out food out of the tank and feed less next time. An anemone that has lost its color still needs food, in small, easily digested pieces. An anemone whose tentacles aren't sticky and won't hold food, or whose mouth is gaping, is showing a sign of trouble (Module 30).

If Then
Food Raw, meaty, smaller than the mouth.
How Flow off, onto the tentacles, never forced.
Spat out hours later Too big, or fed too soon.
Tentacles not sticky, mouth gaping A sign of trouble.

Check your understanding

An anemone under good light is fed a large piece of fish while the pumps run. Half is blown off, a shrimp takes some, and the rest is spat out hours later. What went wrong, and what changes?

Three things: the piece was too big, the flow was on, and nothing kept the shrimp off. Cut pieces smaller than the mouth, turn the flow off, and watch until it's swallowed.

Key point Small pieces, placed, not pushed.

9. Broadcast and target feeding a reef

A reef holds mouths of many sizes. Broadcast feeding has to reach all of them.

Feeding anemones, clams and corals Animal Fed by Anemones Light, plus meaty food it catches Giant clams Light, plus plankton filtered from water Corals Light, plus plankton caught by polyps Broadcast Into the water, for filter feeders Target Placed on one animal Food sizes in a mixed reef Food size Who takes it 2 to 20 microns Phytoplankton Clams, soft corals, sponges, worms 40 to 200 microns Rotifers, fine powders Small-polyp stony corals 1 mm and larger Mysis, cut seafood Large-polyp corals, anemones One size feeds one group A mixed reef needs a mix of sizes Broadcast feeding, step by step Mix the food with tank water Dry powder is wetted first ! Skimmer and filter socks off Or they take the food out Pour into strong flow Keep it moving, not settling Filtration back on After 15 to 30 minutes Feed when the polyps are out For many corals that is after the lights go down

Broadcast feeding puts food into the water for everything that filters or catches it. Target feeding places food on one animal. Corals with large polyps and anemones are target fed pieces they can hold. Corals with small polyps have hundreds of tiny mouths on one colony, so they're fed by broadcast, as are giant clams, sponges and feather worms.

Each group takes a different size. Phytoplankton, at about 2 to 20 microns, feeds giant clams, soft corals, sponges and worms, and it feeds the rotifers and copepods living in the system, which become food in turn. Rotifers and fine powdered foods, roughly 40 to 200 microns, suit small-polyp stony corals. Mysis and chopped seafood, 1 mm and larger, suit large-polyp corals and anemones. A food of one size feeds one group, so a mixed reef is fed a mix of sizes.

To broadcast feed, mix the food with a cup of tank water first; a dry powder that isn't wetted floats or clumps. Turn off the skimmer and take mechanical filtration out of the path, or they remove the food before it's eaten. Pour the mix into strong flow and leave the circulation pumps running so the food stays suspended. Turn the filtration back on after 15 to 30 minutes. Feed when the polyps are out: for many corals that's after the lights go down, and some take food best in the morning. For target feeding, turn the pumps off, put a thicker mix or a piece onto the polyps with a pipette, and turn the pumps back on once it's been taken.

What isn't caught becomes nutrients. Start with a small amount, raise it slowly, and follow nitrate, phosphate and algae as you do (Module 17).

If Then
2 to 20 microns Clams, soft corals, sponges, worms.
40 to 200 microns Small-polyp stony corals.
1 mm and larger Large-polyp corals, anemones.
Skimmer left on The food is removed, not eaten.

Check your understanding

A reef with clams, small-polyp corals and large-polyp corals gets one powdered food each night, with the skimmer running. The large-polyp corals are shrinking and the clams aren't growing. What do you change?

One size feeds one group, and the skimmer is taking much of it. Add phytoplankton for the clams and larger pieces for the large polyps, and turn the skimmer off while feeding.

Key point Mixed reef, mixed sizes.

10. Corals need food as well as light

Light gets most of the attention. Food is the half that's often missed.

Corals need food as well as light Source Gives the coral Light, through the algae in its tissue Energy Food caught by the polyps Energy, plus protein to build new tissue Fed corals grow faster Under the same light: more tissue and more skeleton ! Good light, poor growth Check the feeding before adding more light Match the food to the mouth Some coral mouths are under 0.1 mm across

A coral has two sources of food. The algae living in its tissue use light to make sugars, which give the coral energy. The polyps also catch food from the water, which gives energy and the protein that new tissue is built from. Light alone doesn't supply enough of that building material.

Feeding changes growth. In trials with one stony coral under the same light, fed colonies grew two to eight times more tissue than unfed ones, and built skeleton about 30% faster. Under low light, feeding can keep a coral's skeleton growing. Some corals go further: a Montipora that had lost its algae met all of its daily energy needs by feeding alone. Other corals in the same trial couldn't, so it depends on the coral. A coral that's growing poorly under good light and good water may be short of food, so check the feeding before adding more light.

The food has to fit, and it has to be food the coral can use. The mouth of a flowerpot coral (Goniopora) is only 50 to 100 microns across, so it can't swallow large prey; in a trial it grew best on a very fine, protein-rich prepared food, better than on yeast or phytoplankton. Another coral ate newly hatched Artemia readily and didn't grow any faster. Taking the food isn't proof that it's helping: growth is. Timing can matter too: the flowerpot coral digested best in the morning (Module 27).

If Then
Light Energy.
Food Energy, and protein to build tissue.
Good light, poor growth Check the feeding.

Check your understanding

A coral under good light, in good water, has barely grown in months. It's never fed. What do you check before changing the light?

The feeding. Light gives energy; food gives the protein that new tissue is built from. Feed a food small enough for its mouth, and judge it by growth.

Key point Corals are fed as well as lit.

11. Automatic feeding

On a reef, small food arrives all day. A dosing pump can copy that.

Automatic feeding with a dosing pump Food container Kept cold, kept mixed Dosing pump on a timer Many small doses, day and night Into strong flow Not into the skimmer or a sock ! Warm food spoils Food that needs a refrigerator goes bad in a room-temperature container

Corals and filter feeders do better on many small meals than on one large one, and much of a large one is filtered out before it's eaten. A dosing pump on a timer (Module 19) can add liquid food in small doses through the day and night: phytoplankton, rotifers, or a liquid coral food. Automatic feeders for dry food do the same job for fish.

Food isn't a supplement, and it spoils. A food that's stored in a refrigerator goes bad in a container at room temperature and grows mold. Its container sits in a small refrigerator, or a food made to keep at room temperature is used. Particles settle, so the container is stirred or shaken, or a live culture is kept bubbling with air.

Send the dose into strong flow in the display, not into the skimmer or a filter sock. Smell and look at the food when the container is refilled: food that has gone off is thrown away, not fed.

If Then
Food container Cold and mixed.
Dose Small, often, into strong flow.
Food smells off Throw it away.

Check your understanding

A dosing pump adds a refrigerated coral food from a jar on the stand. The jar smells bad and the corals have stopped putting their polyps out at feeding time. What's wrong?

The food spoiled at room temperature. Throw it away, keep the food cold, or use a food made to keep at room temperature.

Key point Small doses often, from food kept fresh.

12. Live food

Live foods are grown as a chain, each one feeding the next.

The live food chain Phytoplankton Tiny drifting algae Rotifers and copepods Eat the phytoplankton Larvae, small fish, corals Eat the rotifers and copepods ! Keep cultures apart A few rotifers can eat a whole phytoplankton culture ! Keep a backup Cultures crash without warning

Phytoplankton is tiny drifting algae. Rotifers and copepods eat phytoplankton, and larvae, small fish and corals eat the rotifers and copepods. Newly hatched Artemia, or brine shrimp, are a larger live food for bigger larvae and small fish. Fish larvae often can't be raised without live food: it's small enough, and its movement makes them strike.

Cultures are kept clean and apart. A few rotifers carried into a phytoplankton culture will eat it, so each culture has its own tools and phytoplankton is handled first. Cultures can crash without warning, so a backup of each is kept running.

If Then
Rotifers in the phytoplankton The culture gets eaten.
One culture, no backup A crash leaves nothing.

Check your understanding

A phytoplankton culture turns clear over two days, and rotifers are swimming in it. What happened, and how is it prevented?

Rotifers got in and ate it. Each culture has its own tools, phytoplankton is handled first, and a backup is kept.

Key point Clean, apart, backed up.

13. Enrichment and storage

Food is only as good as what's in it when it's eaten.

Enriching live food Live prey Rotifers or Artemia Enrich Feed them DHA and EPA Feed out soon Within hours ! It fades A day after enriching, the prey is no longer enriched Storing food Food Keep it Dry food Cool, dry, sealed, away from the tank Opened dry food Replace within 3 to 6 months Frozen food Frozen: never thawed and refrozen ! Vitamins go first Old food can look fine and feed poorly

Live prey is only as nutritious as what it has eaten. Rotifers and Artemia are low in the fatty acids DHA and EPA that marine fish larvae need, so before they're fed out they're enriched: fed a food rich in those fatty acids. Enrichment fades as the prey uses it up; within hours it drops, and a day later the prey is no longer enriched. Enriched prey is fed out soon.

Stored food loses value too, vitamins first. Dry food kept beside a warm, humid tank goes off fastest: keep it cool, dry and sealed, away from the tank, and replace it within 3 to 6 months of opening. Frozen food stays frozen until it's used, and is never thawed and refrozen. Old food can look fine and still feed poorly.

If Then
Rotifers enriched yesterday No longer enriched.
Dry food by the warm tank Loses vitamins fast.

Check your understanding

Rotifers were enriched yesterday morning and are fed to larvae this morning. What's the problem?

A day later they're no longer enriched. Enrich, then feed out within hours.

Key point Fresh food, fed soon.

14. When feeding goes wrong

Feeding problems are often a combination. Check everything before fixing anything.

When feeding goes wrong You see Check Algae spreading, food on the sand How much goes in One thin fish, others fine Who gets the food, and its size Thin grazer The type of food All fish dull, slow growth Food age and variety Larvae failing Prey size and enrichment timing ! Often more than one Check them all before fixing

Work from what you see. Algae spreading with food on the sand points to the amount. One thin fish among healthy ones points to who's getting the food, or its size. A thin grazer points to the type of food. A whole tank that's dull and growing slowly points to old food or too little variety. Larvae that fail point to prey size and enrichment timing.

More than one of these is often true at once: the food is stale and there's too much of it; the pellet is too big and the bold fish takes it first. Check them all, then fix each cause.

Feeding drill: on the course website this is an interactive drill. Read what you see, check each part, and fix every cause.

If Then
One thin fish Who eats, and food size.
Algae and dull fish Amount, and food age.

Check your understanding

One small fish is thin while the others are fat, and there's food on the sand every day. How many problems is that?

At least two: that fish isn't getting food, or can't eat its size, and too much is going in overall.

Key point Check all, then fix each.

15. What you see, and what it means

Food on the sand after meals Too much is going in.
Algae spreading Excess nutrients, often overfeeding.
Thin fish, pinched belly Not enough food reaching it.
Takes food and spits it out Too big, wrong food, or sick.
Thin grazer Needs seaweed, through the day.
Fish ignores food Often the first sign of trouble.
Anemone spits food out hours later Piece too big, or fed too soon.
Anemone won't hold food A sign of trouble.
Coral not growing under good light It may be short of food.
Liquid food smells off It spoiled. Throw it away and keep it cold.
Phytoplankton culture clearing Rotifers got in.
Larvae failing on rotifers Check enrichment timing and prey size.
All fish dull Old food or too little variety.

Module 26 test

30 questions. Pass with 27 or more. Opens after you pass every section check in this module. Sign in to start.

Sources

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  3. RateMyFishTank: Tips for feeding saltwater aquarium fish
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  10. Science Insights: Does fish food go bad?
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  20. Gensou: Broadcast feeding SPS colonies technique guide
  21. Reef Builders: How to make a Wandell auto refrigerated feeder
  22. Nano-Reef: Dosing coral feed (food spoiling in a dosing container)
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  27. Ferrier-Pages, Hoogenboom and Houlbreque (2011): The role of plankton in coral trophodynamics
  28. Michael Maddox, President of IMARCS: coral feeding gets too little attention next to light, feeding a range of particle sizes to support every level of the food web, nutrients as food at the right concentration, resting clownfish that spring out to eat, problems as combinations